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锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟节硷拷锟矫达拷纽锟斤拷锟斤拷锟斤拷锟斤拷拇锟斤拷锟斤拷锟斤拷锟斤拷锟轿拷锟斤拷鼐锟斤拷锟斤拷峁╋拷舜锟斤拷锟斤拷锟绞筹拷锟斤拷锟斤拷锟诫。
锟斤拷图片锟结供:John Eastcott and Yva Momatiuk/National Geographic/Getty Images锟斤拷
一锟斤拷锟铰碉拷锟叫撅拷锟阶达拷预锟解，锟斤拷锟斤拷锟斤拷锟角诧拷取锟斤拷锟斤拷锟斤拷预锟斤拷锟斤拷施锟斤拷锟斤拷锟斤拷息锟节硷拷锟矫达拷纽锟斤拷锟斤拷锟斤拷锟斤拷锟侥达拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷群锟斤拷锟角斤拷锟斤拷未锟斤拷20锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟矫达拷锟斤拷锟斤拷锟斤拷锟剿癸拷写锟斤拷锟斤拷锟斤拷锟斤拷学锟斤拷锟斤拷业锟斤拷锟斤拷学锟斤拷Jeffrey Hutchings锟斤拷示锟斤拷“锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟侥癸拷锟斤拷一锟街猴拷锟斤拷锟斤拷锟斤拷锟斤拷群锟斤拷锟筋骇锟斤拷锟斤拷锟脚猴拷锟斤拷锟剿凤拷锟秸碉拷锟斤拷息锟斤拷”
锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷业锟皆诧拷锟姐繁锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷循锟斤拷锟斤拷一锟斤拷锟斤拷锟脚★拷锟斤拷20锟斤拷锟酵猴拷锟节筹拷锟斤拷50锟斤拷拇锟斤拷锟斤拷锟斤拷毯螅锟斤拷么锟斤拷锟斤拷锟斤拷水锟斤拷业锟斤拷锟斤拷锟斤拷锟斤拷90锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟揭碉拷锟斤拷锟斤拷锟斤拷锟斤拷婕达拷锟教ㄒ幌碉拷薪锟斤拷睿拷锟斤拷诖锟斤拷芄锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟饺猴拷锟斤拷锟斤拷母锟斤拷铡锟饺伙拷锟斤拷锟斤拷锟�15锟斤拷锟斤拷锟斤拷薏锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟襟，碉拷锟截碉拷锟斤拷锟斤拷锟侥ｏ拷锟矫伙拷斜锟斤拷殖锟斤拷锟斤拷锟斤拷募锟斤拷锟绞碉拷锟斤拷希锟斤拷锟叫╋拷锟斤拷锟斤拷锟斤拷群锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟铰斤拷锟斤拷锟叫撅拷锟斤拷员锟斤拷锟斤拷锟铰筹拷锟斤拷摹锟斤拷锟斤拷么锟斤拷锟揭碉拷锟剿拷锟斤拷锟窖э拷锟街撅拷锟斤拷媳锟斤拷锟斤拷锟斤拷锟揭伙拷芯锟斤拷晒锟斤拷锟�
锟斤拷锟矫达拷锟斤拷业锟诫海锟襟部碉拷锟斤拷锟斤拷学锟斤拷Douglas Swain锟斤拷Ghislain Chouinard锟斤拷锟斤拷锟矫斤拷锟斤拷锟斤拷锟狡碉拷锟斤拷业锟斤拷锟斤拷模锟酵ｏ拷锟斤拷锟斤拷锟斤拷锟斤拷纽锟斤拷锟斤拷锟斤拷锟较诧拷圣锟斤拷锟斤拷斯锟斤拷锟饺拷锟斤拷锟侥达拷锟斤拷锟斤拷锟斤拷群锟斤拷未锟斤拷锟斤拷锟斤拷锟斤拷预锟解。锟斤拷些模锟酵匡拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷群锟侥凤拷殖锟斤拷——锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷群锟叫的憋拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷状锟斤拷锟皆硷拷锟斤拷然锟斤拷锟斤拷锟绞★拷锟斤拷锟秸碉拷预锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷思——锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷薏锟斤拷蹋锟斤拷锟斤拷锟斤拷诤锟斤拷锟斤拷喜锟斤拷锟斤拷锟斤拷锟斤拷锟饺猴拷锟斤拷锟轿达拷锟�20锟斤拷锟斤拷锟斤拷锟斤拷锟饺伙拷锟斤拷锟斤拷悴讹拷锟斤拷锟斤拷锟斤拷锟斤拷业锟筋动锟斤拷锟斤拷全锟斤拷止锟斤拷锟斤拷些锟斤拷群也锟斤拷锟斤拷38锟斤拷锟斤拷锟斤拷失锟斤拷锟斤拷锟斤拷
Swain锟斤拷Chouinard锟斤拷为锟斤拷锟斤拷锟斤拷锟揭伙拷锟斤拷锟斤拷锟斤拷要锟斤拷锟斤拷锟斤拷锟节斤拷锟斤拷锟斤拷锟叫ｏ拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷寻锟斤拷锟截高★拷没锟斤拷锟斤拷知锟斤拷锟斤拷锟斤拷为什么锟斤拷锟斤拷Swain锟狡诧拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷诤锟斤拷锟斤拷牟锟绞筹拷锟轿拷泳锟斤拷锟斤拷隆锟饺伙拷锟斤拷锟斤拷饣乖讹拷锟街癸拷诖锟�——锟斤拷锟捷硷拷锟矫达拷锟窖э拷锟斤拷锟�2003锟疥公锟斤拷锟斤拷一锟斤拷锟斤拷锟捷ｏ拷锟斤拷锟斤拷Scotian锟斤拷陆锟杰碉拷锟斤拷锟斤拷锟斤拷群锟斤拷锟斤拷也锟斤拷锟斤拷锟斤拷锟斤拷90锟斤拷锟斤拷锟斤拷锟斤拷锟劫★拷锟斤拷锟解，Swain指锟斤拷锟斤拷锟斤拷然锟斤拷锟斤拷锟节硷拷锟矫达拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷群锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷茫锟斤拷锟斤拷锟斤拷锟斤拷强锟斤拷锟揭诧拷锟斤拷锟轿达拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟酵拷锟斤拷锟斤拷锟斤拷狻�
Hutchings强锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷学锟揭达拷统锟较硷拷锟借，锟斤拷锟酵Ｖ癸拷锟揭碉拷锟斤拷蹋锟斤拷锟斤拷锟斤拷锟饺猴拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟街凤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟叫撅拷“却锟角斤拷锟斤拷锟街硷拷锟斤拷锟轿拷锟斤拷鄣锟揭伙拷锟斤拷浅锟斤拷锟斤拷锟阶⒛匡拷锟斤拷锟街�”锟斤拷谈锟斤拷一锟斤拷锟斤拷锟斤拷锟斤拷群锟斤拷锟斤拷锟斤拷锟紿utchings说锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷之前锟斤拷锟斤拷锟斤拷未锟斤拷锟角癸拷锟斤拷锟街匡拷锟斤拷锟皆★拷然锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷Woods Hole锟侥癸拷锟揭猴拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷值锟斤拷锟揭碉拷锟斤拷锟窖э拷锟絉alph Mayo锟斤拷为锟斤拷锟斤拷锟街观碉拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷沙锟睫的斤拷小锟斤拷锟斤拷锟斤拷锟斤拷群锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟侥★拷锟斤拷说锟斤拷“锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟饺猴拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟�”锟斤拷然锟斤拷锟斤拷锟斤拷诩锟斤拷么锟斤拷锟斤拷锟剿碉拷锟斤拷锟斤拷锟揭伙拷锟叫⌒★拷陌锟轿匡拷锟斤拷锟斤拷锟皆达拷锟斤拷锟窖憋拷锟� 群锟斤拷锟斤拷
锟斤拷锟斤拷锟斤拷锟矫达拷锟斤拷业锟斤拷水锟斤拷锟斤拷学锟斤拷志锟斤拷锟斤拷
Canadian Journal of Fisheries and Aquatic Sciences锟斤拷锟斤拷65(11): 2315–2319锟斤拷Douglas P. Swain锟斤拷Ghislain A. Chouinard锟斤拷
