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| 锟斤拷趾锟斤拷锟斤拷为锟斤拷锟杰斤拷锟斤拷锟斤拷锟斤拷 |
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锟斤拷锟斤拷挪锟斤拷锟斤拷锟狡�
锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟轿拷锟矫伙拷锟斤拷锟斤拷锟阶ワ拷盏某锟斤拷锟街猴拷锟斤拷佣锟斤拷锟剿拷锟揭诧拷弑锟剿拷值墓锟斤拷埽锟斤拷锟矫达拷锟斤拷碌目锟窖э拷芯锟斤拷锟斤拷锟斤拷锟斤拷锟金案ｏ拷锟街讹拷锟教的斤拷趾锟斤拷锟斤拷锟斤拷为锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟狡的★拷
锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷学锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟街猴拷榷探锟街猴拷姆迅锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟金动ｏ拷锟斤拷也锟角帮拷锟斤拷锟斤拷住锟节达拷锟皆拷锟斤拷锟斤拷锟斤拷锟斤拷锟阶凤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟街伙拷锟�
锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟街碉拷锟斤拷要锟斤拷志
锟斤拷锟矫大卡讹拷锟斤拷锟斤拷锟窖э拷锟斤拷锟窖э拷铱锟斤拷锟斤拷锟�·锟斤拷锟斤拷锟斤拷(Campbell Rolian)说锟斤拷“锟斤拷榷探锟街猴拷锟斤拷锟斤拷锟街猴拷锟斤拷硕锟绞憋拷锟揭拷锟斤拷猓拷锟揭拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷鼙锟斤拷锟斤拷锟斤拷锟斤拷榷锟斤拷锟街灰拷锟斤拷谴锟斤拷麓锟斤拷锟斤拷芏锟斤拷锟斤拷锟矫达拷锟饺谎★拷锟斤拷锟斤拷锟斤拷锟斤拷诙探锟街猴拷锟斤拷恕锟�”锟斤拷锟斤拷锟斤拷槌わ拷喽拷铮拷锟斤拷锟斤拷锟斤拷锟斤拷枪锟较碉拷锟斤拷锟侥猴拷锟斤拷锟缴ｏ拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟街猴拷锟斤拷锟斤拷锟斤拷锟斤拷锟侥斤拷趾锟斤拷锟斤拷锟斤拷锟街菏拷侄锟叫★拷锟斤拷芯锟矫伙拷锟斤拷锟饺ｏ拷只锟斤拷锟届开锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟杰的讹拷锟斤拷慕锟街和拷锟斤拷芏獭锟矫拷凸锟斤拷锟揭恍╋拷锟斤拷锟斤拷爪锟接硷拷锟斤拷锟斤拷全锟缴斤拷锟狡伙拷锟斤拷锟狡癸拷锟缴★拷锟斤拷锟斤拷锟斤拷锟斤拷锟叫撅拷小锟斤拷锟缴达拷锟斤拷知锟斤拷锟斤拷锟斤拷锟斤拷锟角凤拷锟斤拷越锟斤拷锟斤拷私诺锟斤拷锟斤拷锟斤拷锟斤拷臁�
锟斤拷然锟斤拷锟斤拷锟杰讹拷锟斤拷锟斤拷锟斤拷锟剿碉拷锟斤拷要锟皆很猴拷锟狡测。锟斤拷锟角ｏ拷锟斤拷锟筋技锟杰讹拷锟斤拷锟斤拷锟斤拷锟饺肥碉拷锟斤拷胁锟酵帮拷锟斤拷锟斤拷锟斤拷澹猴拷锟斤拷锟斤拷锟斤拷锟斤拷猓伙拷屑锟斤拷锟斤拷锟斤拷锟斤拷锟竭憋拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷没锟斤拷一锟街讹拷锟斤拷锟斤拷锟斤拷诔锟斤拷鹊锟斤拷锟斤拷锟斤拷鲁锟绞憋拷浔硷拷堋锟斤拷锟斤拷锟斤拷呛锟斤拷喙凤拷锟斤拷锟斤拷锟街伙拷锟斤拷诤锟斤拷锟斤拷锟斤拷锟斤拷锟狡伙拷时锟街筹拷途锟斤拷袭锟斤拷锟皆ｏ拷锟节革拷锟斤拷锟斤拷锟斤拷锟斤拷锟缴ナэ拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟揭拷锟斤拷锟斤拷谋锟斤拷锟斤拷锟斤拷墙锟斤拷锟斤拷锟斤拷锟斤拷锟酵拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟揭伙拷锟斤拷锟揭拷锟街撅拷锟�
锟斤拷锟斤拷锟斤拷锟斤拷锟叫撅拷锟斤拷锟斤拷一位锟斤拷员锟斤拷锟斤拷锟斤拷锟窖э拷锟斤拷锟窖э拷业锟斤拷锟斤拷·锟斤拷锟斤拷锟斤拷(Daniel Lieberman)锟斤拷锟杰ｏ拷锟斤拷锟叫关达拷锟皆拷某锟酵撅拷锟斤拷锟斤拷锟斤拷锟斤拷希锟斤拷执锟斤拷锟斤拷锟结构锟侥很讹拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷要锟斤拷锟矫★拷锟斤拷锟斤拷帽鹊锟斤拷桑锟斤拷锟斤拷锟斤拷鸬奖锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷茫锟斤拷锟街拷锟斤拷谐锟斤拷锟截节ｏ拷锟斤拷锟轿诧拷锟斤拷锟斤拷锟斤拷锟角憋拷锟斤拷锟饺讹拷锟斤拷锟斤拷锟斤拷锟斤拷锟竭★拷同锟斤拷锟斤拷锟斤拷锟皆对憋拷锟杰活动锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷械锟斤拷锟斤拷卸取锟�
锟斤拷趾锟斤拷锟斤拷锟接对硷拷锟斤拷锟斤拷锟斤拷
锟斤拷趾锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷一锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟剿碉拷锟�“锟斤拷锟斤拷浅锟斤拷屎锟斤拷锟斤拷锟斤拷埽锟斤拷锟揭伙拷锟斤拷潭锟斤拷锟绞癸拷锟斤拷锟斤拷锟戒化锟斤拷锟斤拷锟斤拷锟藉。锟斤拷锟角诧拷锟斤拷锟角筹拷色锟侥讹拷锟斤拷锟剿讹拷员锟斤拷锟斤拷锟揭伙拷锟角碉拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷某锟斤拷锟窖★拷帧锟�”锟斤拷锟斤拷锟斤拷锟斤拷锟矫筹拷锟节筹拷锟杰碉拷锟桔讹拷锟斤拷未锟斤拷锟秸憋拷锟斤拷堋锟斤拷锟斤拷锟斤拷锟剿癸拷锟斤拷锟斤拷荽锟窖э拷锟斤拷锟斤拷锟窖э拷锟皆硷拷锟�·锟斤拷锟斤拷斯(John Hawks)说锟斤拷“锟竭猴拷锟斤拷锟斤拷锟斤拷锟斤拷一锟斤拷锟斤拷锟斤拷锟藉部位锟斤拷锟斤拷锟斤拷说锟斤拷些锟斤拷锟藉构锟斤拷锟斤拷为锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟狡ｏ拷锟斤拷为确锟叫碉拷说锟斤拷锟斤拷为锟斤拷锟杰达拷锟斤拷摹锟�”锟斤拷锟斤拷斯没锟叫参硷拷锟斤拷锟斤拷锟斤拷锟斤拷锟叫撅拷锟斤拷
锟斤拷锟角ｏ拷锟斤拷锟斤拷锟斤拷锟斤拷锟叫撅拷锟斤拷锟斤拷证锟斤拷锟剿斤拷趾锟斤拷锟杰诧拷锟斤拷锟斤拷要锟皆★拷锟斤拷锟斤拷锟叫撅拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷一锟节★拷实锟斤拷锟斤拷锟斤拷学锟斤拷志锟斤拷锟斤拷Journal of Experimental Biology锟斤拷锟较★拷锟斤拷锟斤拷锟斤拷前锟竭伙拷锟斤拷时锟斤拷一只锟斤拷锟节匡拷锟叫ｏ拷锟斤拷一只锟斤拷锟节碉拷锟芥，锟斤拷锟斤拷一锟诫到锟侥凤拷之锟斤拷锟斤拷锟斤拷锟斤拷恰锟斤拷锟斤拷锟斤拷前锟斤拷锟较★拷锟斤拷锟斤拷锟斤拷说锟斤拷“锟斤拷锟竭诧拷时锟斤拷一只锟斤拷锟斤拷锟斤拷锟斤拷一锟斤拷前锟斤拷锟斤拷一只锟斤拷锟窖撅拷锟脚地★拷锟斤拷锟窖撅拷转锟斤拷锟剿诧拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷慕锟街猴拷锟斤拷锟斤拷锟斤拷芏锟斤拷锟斤拷锟斤拷锟斤拷锟缴革拷锟斤拷墓锟斤拷锟斤拷锟斤拷贫锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷前锟斤拷”
实锟斤拷锟叫ｏ拷15锟斤拷志愿锟斤拷锟斤拷一锟斤拷锟斤拷压锟斤拷十锟斤拷锟斤拷锟叫的憋拷锟斤拷锟斤拷一锟斤拷锟斤拷埽锟揭伙拷锟斤拷锟竭ｏ拷锟斤拷锟斤拷锟斤拷锟斤拷锟叫撅拷小锟斤拷锟斤拷锟斤拷歉锟斤拷锟斤拷锟斤拷锟斤拷施锟接碉拷锟斤拷锟斤拷锟斤拷锟斤拷锟剿凤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷纸锟街猴拷锟斤拷锟街灰拷锟斤拷锟�20%锟斤拷锟斤拷锟斤拷锟侥电动锟斤拷(motor force)却锟斤拷原锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟角匡拷锟皆从革拷为直锟桔碉拷锟斤拷锟轿帮拷疃拷锟斤拷锟斤拷锟斤拷锟揭伙拷锟斤拷猓貉癸拷锟斤拷锟街э拷锟街拷锟侥撅拷锟斤拷锟绞癸拷芨锟斤拷锟斤拷锟斤拷锟�
锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟街ｏ拷锟斤拷锟斤拷趾锟斤拷锟斤拷锟斤拷锟斤拷停止锟筋动锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷去锟斤拷锟斤拷锟杰猴拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟角帮拷锟侥活动时锟斤拷锟斤拷要锟侥费革拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷沙锟斤拷锟街猴拷姆训模锟斤拷锟斤拷锟斤拷锟斤拷锟剿对硷拷锟斤拷锟窖癸拷锟斤拷锟斤拷锟斤拷耍锟斤拷锟斤拷锟绞癸拷锟斤拷为锟斤拷然选锟斤拷锟斤拷锟斤拷锟狡凤拷锟�
凭锟斤拷锟斤拷锟斤拷锟斤拷杀锟斤拷锟酵讹拷锟斤拷
锟斤拷石锟斤拷录也锟斤拷锟斤拷锟结供锟剿猴拷锟绞碉拷锟斤拷证锟斤拷锟斤拷锟斤拷猿锟侥斤拷趾锟斤拷锟较凤拷锟斤拷猿(锟斤拷锟斤拷锟斤拷锟斤拷锟皆硷拷锟斤拷锟�)锟斤拷趾锟斤拷锟斤拷锟斤拷锟较凤拷锟斤拷猿锟侥斤拷趾锟街憋拷锟斤拷锟斤拷锟侥斤拷趾锟斤拷锟斤拷锟街达拷锟剿撅拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷斯指锟斤拷锟斤拷锟斤拷途锟斤拷锟斤拷锟斤拷锟窖硷拷为锟斤拷锟斤拷锟斤拷“锟斤拷锟斤拷锟斤拷诘牡胤锟斤拷锟斤拷锟斤拷煞浅锟斤拷锟斤拷拥锟斤拷幕锟斤拷锟接χэ拷锟�(cultural adaptation)锟侥ｏ拷锟斤拷锟斤拷锟斤拷锟劫★拷水锟斤拷源锟斤拷锟芥，锟斤拷锟斤拷锟斤拷锟剿回家等等★拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟角凤拷锟斤拷锟斤拷锟叫╋拷幕锟斤拷锟接χわ拷荩锟斤拷纸锥渭锟斤拷锟矫伙拷校锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟节★拷”
然锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷指锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷然锟皆达拷锟酵讹拷锟斤拷为食锟斤拷锟斤拷锟斤拷锟斤拷杀锟斤拷锟酵讹拷锟斤拷锟斤拷锟斤拷锟斤拷锟酵讹拷锟斤拷锟街伙拷锟斤拷诰锟斤拷千锟斤拷前锟斤拷锟斤拷锟斤拷锟斤拷锟侥★拷锟斤拷锟斤拷锟斤拷说锟斤拷“锟斤拷锟角碉拷锟斤拷锟饺ｏ拷锟斤拷些锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟介长锟洁动锟斤，锟斤拷锟斤拷锟缴憋拷锟斤拷投锟斤拷铮匡拷锟斤拷锟斤拷锟斤拷强锟斤拷锟阶凤拷锟斤拷恰锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷欠杀锟斤拷锟斤拷锟斤拷遣锟斤拷锟揭伙拷弑锟斤拷埽锟揭伙拷叽锟斤拷锟斤拷锟斤拷锟斤拷遣锟斤拷锟狡撅拷俣锟斤拷峡锟斤拷锟斤拷锟侥ｏ拷锟斤拷锟斤拷凭锟斤拷锟斤拷锟斤拷锟斤拷”
锟斤拷然锟斤拷锟斤拷锟斤拷锟斤拷痰锟酵诧拷锟斤拷锟斤拷锟斤拷锟斤拷锟窖憋拷锟揭伙拷锟斤拷锟斤拷锟斤拷锟斤拷只疃拷锟斤拷锟接诧拷锟叫拷锟斤拷锟斤拷顺锟斤拷锟斤拷锟斤拷锟斤拷艿牟锟斤拷锟斤拷鸲锟斤拷庸糯锟斤拷锟斤拷锟斤拷锟窖癸拷锟斤拷头懦锟斤拷锟斤拷锟斤拷锟斤拷堑锟剿拷锟斤拷只锟斤拷锟斤拷锟绞裁达拷锟斤拷锟斤拷兀锟斤拷锟斤拷锟斤拷锟剿碉拷锟斤拷锟斤拷锟剿狄伙拷卸锟斤拷锟斤拷岱拷锟斤拷锟轿憋拷锟斤拷纾�“锟斤拷锟斤拷锟斤拷一锟斤拷锟叫癸拷锟斤拷锟藉构锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷毡锟斤拷锟斤拷狻ｏ拷锟轿拷锟斤拷遣锟斤拷锟斤拷锟揭拷探锟街猴拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟矫ｏ拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷锟斤拷谈锟桔短斤拷趾锟斤拷锟斤拷锟角凤拷锟斤拷锟绞э拷锟�” 锟斤拷锟斤拷源锟斤拷 锟斤拷锟剿科硷拷 孝锟侥ｏ拷
锟斤拷锟斤拷实锟斤拷锟斤拷锟斤拷学锟斤拷志锟斤拷锟斤拷
Journal of Experimental Biology锟斤拷锟斤拷doi: 10.1242/jeb.019885 锟斤拷Campbell Rolian, William Werbel锟斤拷
